Chapter 11 Organisms and Populations
Introduction
Levels of Biological Organization:
- Macromolecules
- Cells
- Tissues
- Organs
- Individual organisms
- Population
- Communities
- Ecosystems
- Biomes
Types of Questions:
- “How-type” Questions: Seek the mechanism behind a process.
- “Why-type” Questions: Seek the significance of a process.
Example with a Bulbul Bird:
- How does the bird sing? (Mechanism: Operation of voice box, vibrating bone)
- Why does the bird sing? (Significance: Communication with mate during breeding season)
Observing Nature with a Scientific Mindset:
- Encourages asking interesting questions (e.g., Why are night-blooming flowers white? How does a bee locate nectar? Why does a cactus have thorns? How does a chick recognize its mother?)
Ecology:
- Studies interactions among organisms and between organisms and their physical (abiotic) environment.
- Concerned with four levels of biological organization: organisms, populations, communities, and biomes.
Focus in This Chapter:
- Exploring ecology at the population level.
11.1 POPULATIONS
11.1.1 Population Attributes
Definition of Population:
- A population consists of individuals of the same species living in a defined area, sharing or competing for resources, and potentially interbreeding.
Examples of Populations:
- Cormorants in a wetland, rats in a dwelling, teakwood trees in a forest, bacteria in a culture plate, lotus plants in a pond.
Population Ecology’s Importance:
- Natural selection operates at the population level to evolve desired traits, linking ecology to population genetics and evolution.
Population Attributes:
- Birth and Death Rates: Expressed as per capita births and deaths, indicating change in numbers relative to the population size.
- Sex Ratio: Ratio of males to females in a population.
- Age Structure: Distribution of individuals across different age groups, represented in an age pyramid.
- Population Size and Density: Represented by the total number of individuals per unit area or volume, but sometimes measured more meaningfully in terms of biomass or cover.
Population Size Measurement:
- Can be challenging in large or inaccessible populations, requiring indirect estimation methods (e.g., tiger census based on pug marks and fecal pellets).
Importance of Population Size in Ecology:
- Essential for studying ecological processes such as competition, predation, and pesticide impacts, as these processes often affect population size.
11.1.2 Population Growth
Population Dynamics:
- Population size changes over time due to factors like food availability, predation, weather, etc.
- Changes in population density indicate whether a population is flourishing or declining.
Four Basic Processes Affecting Population Density:
- Natality (births)
- Mortality (deaths)
- Immigration (incoming individuals)
- Emigration (outgoing individuals)
Population Density Formula:
- 𝑁𝑡+1=𝑁𝑡+[(𝐵+𝐼)−(𝐷+𝐸)]Nt+1=Nt+[(B+I)−(D+E)]
- Population density at time t + 1 equals density at time t plus net births and immigration minus deaths and emigration.
Factors Influencing Population Density:
- Births and deaths are usually the most significant factors affecting population density.
- Immigration and emigration become important under special conditions (e.g., colonization of a new habitat).
Population Growth Models:
- Exponential Growth: Occurs when resources are unlimited, leading to unimpeded population growth.
- Population growth rate (𝑑𝑁/𝑑𝑡dN/dt) equals the intrinsic rate of natural increase (𝑟r) times the population size (𝑁N): 𝑑𝑁/𝑑𝑡=𝑟𝑁dN/dt=rN.
- Results in a J-shaped curve when plotted over time.
- Formula for exponential growth: 𝑁𝑡=𝑁0⋅𝑒𝑟𝑡Nt=N0⋅ert, where 𝑒e is the base of natural logarithms (2.71828).
Intrinsic Rate of Natural Increase (r):
- Represents the potential growth rate of a population under ideal conditions.
- Varies among species and can be used to assess impacts of factors on population growth.
Exponential Growth:
- Occurs when resources are unlimited, leading to unimpeded population growth.
- Population size increases rapidly and continuously, following an exponential curve (J-shaped curve).
Anecdote of Chessboard and Wheat Grains:
- Demonstrates the dramatic effect of exponential growth.
- Even a seemingly small initial quantity can lead to an enormous population size under exponential growth.
Logistic Growth:
- Reflects the realistic growth pattern of populations with limited resources.
- Includes phases such as lag phase, acceleration, deceleration, and reaching an asymptote (carrying capacity).
- Described by the Verhulst-Pearl Logistic Growth equation:
$$\frac{dN}{dt}=rN(1-\frac{N}{K})$$
- 𝑁N: Population density at time t
- 𝑟r: Intrinsic rate of natural increase
- 𝐾K: Carrying capacity (maximum population size that the habitat can support)
Population Growth in Nature:
- No population in nature experiences unlimited resources, so logistic growth is more common and realistic.
- Governments recognize the need to control human population growth due to limited resources and competition.
Applying Growth Models to Census Data:
- Use government census data for India over the last 100 years to analyze population growth patterns.
- Plot the population figures over time to identify whether exponential or logistic growth patterns are evident.11.1.3 Life History Variation.