Pteridophytes include horsetails and ferns, and they are utilized for medicinal purposes, soil-binding, and ornamental gardening.
Evolutionarily, they are the first terrestrial plants to possess vascular tissues, namely xylem and phloem.
Habitat:
Pteridophytes are commonly found in cool, damp, and shady places, although some species may thrive in sandy-soil conditions.
Life Cycle:
In contrast to bryophytes, where the dominant phase is the gametophyte, the main plant body in pteridophytes is the sporophyte.
The sporophyte is differentiated into true roots, stems, and leaves, each possessing well-differentiated vascular tissues.
Leaves in pteridophytes may be small (microphylls) as in Selaginella or large (macrophylls) as in ferns.
Sporangia, which produce spores by meiosis, are borne on leaf-like appendages called sporophylls. In some cases, sporophylls may form compact structures called strobili or cones.
Spores germinate to produce inconspicuous, small but multicellular, free-living, mostly photosynthetic thalloid gametophytes called prothalli.
Reproduction:
Gametophytes bear male and female sex organs called antheridia and archegonia, respectively.
Water is necessary for the transfer of male gametes (antherozoids) to the archegonium for fertilization, resulting in the formation of a zygote.
The zygote develops into a multicellular, well-differentiated sporophyte, which is the dominant phase of the pteridophytes.
Spore Types:
Most pteridophytes are homosporous, producing spores of similar kinds.
However, genera like Selaginella and Salvinia are heterosporous, producing two kinds of spores: macrospores (large) and microspores (small).
Classification:
Pteridophytes are further classified into four classes: Psilopsida (Psilotum); Lycopsida (Selaginella, Lycopodium); Sphenopsida (Equisetum); and Pteropsida (Dryopteris, Pteris, Adiantum).
3.4 GYMNOSPERMS
Definition:
Gymnosperms are plants in which the ovules are not enclosed by any ovary wall, remaining exposed both before and after fertilization. Hence, the seeds are naked.
Description:
Gymnosperms include medium-sized trees or tall trees and shrubs. For instance, the giant redwood tree Sequoia is one of the tallest tree species.
Their roots are generally taproots, and some genera have fungal associations with mycorrhiza or coralloid roots associated with N2-fixing cyanobacteria.
Stems may be unbranched (as in Cycas) or branched (as in Pinus and Cedrus).
Leaves may be simple or compound, with adaptations to withstand extremes of temperature, humidity, and wind. For example, conifers have needle-like leaves with a thick cuticle and sunken stomata to reduce water loss.
Reproduction:
Gymnosperms are heterosporous, producing haploid microspores and megaspores.
Microspores develop into pollen grains, which are highly reduced male gametophytes confined to a limited number of cells.
Macrospores develop into multicellular female gametophytes within the ovules, each bearing two or more archegonia or female sex organs.
Unlike bryophytes and pteridophytes, the male and female gametophytes in gymnosperms do not have an independent free-living existence. They remain within the sporangia retained on the sporophytes.
Pollination and Fertilization:
Pollen grains are released from the microsporangium and carried by air currents to the opening of ovules borne on megasporophylls.
The pollen tube carrying the male gametes grows towards the archegonia in the ovules and discharges its contents near the mouth of the archegonia.
Following fertilization, the zygote develops into an embryo, and the ovules develop into seeds, which remain uncovered.
3.5 ANGIOSPERMS
Definition:
Angiosperms, also known as flowering plants, are characterized by the development of specialized structures called flowers, within which pollen grains and ovules are produced.
Seed Enclosure:
Unlike gymnosperms where ovules are naked, seeds in angiosperms are enclosed within fruits, which develop from the fertilized ovary of the flower.
Habitat and Diversity:
Angiosperms are an exceptionally large group of plants found in a wide range of habitats.
They vary greatly in size, ranging from tiny plants like Wolffia to towering trees like Eucalyptus, which can grow over 100 meters tall.
Importance:
Angiosperms are of immense economic and ecological importance, providing food, fodder, fuel, medicines, and numerous other commercially significant products.
Classification:
Angiosperms are divided into two main classes: dicotyledons (dicots) and monocotyledons (monocots).
Dicots typically have seeds with two cotyledons, netted venation in leaves, and flower parts in multiples of four or five.
Monocots usually have seeds with a single cotyledon, parallel venation in leaves, and flower parts in multiples of three.