4.2.7 Phylum – Arthropod
Huge Diversity:
- Arthropods constitute the largest phylum in the animal kingdom, comprising over two-thirds of all named species on Earth.
- They exhibit remarkable diversity in terms of morphology, habitat, and behavior.
Organ-System Level of Organization:
- Arthropods have organ-system level of organization, with distinct organ systems performing specific functions.
Bilateral Symmetry:
- They display bilateral symmetry, with a distinct left and right side.
Triploblastic and Coelomate:
- Arthropods are triploblastic organisms, consisting of three embryonic germ layers: ectoderm, mesoderm, and endoderm.
- They are coelomate animals, possessing a true body cavity (coelom) that is completely lined with mesoderm.
Chitinous Exoskeleton:
- The body of arthropods is covered by a tough, chitinous exoskeleton, providing support and protection.
- This exoskeleton is molted periodically to accommodate growth.
Body Segmentation:
- Arthropods typically have a body divided into distinct regions: head, thorax, and abdomen.
Jointed Appendages:
- Arthropods are characterized by jointed appendages (arthros-jointed, poda-appendages), which are used for locomotion, feeding, and sensory perception.
Respiratory Organs:
- Arthropods may have various respiratory organs, including gills, book gills, book lungs, or a tracheal system, depending on the species.
Circulatory System:
- The circulatory system of arthropods is of the open type, where blood is pumped into the body cavity and bathes the organs directly.
Sensory Organs:
- Arthropods possess sensory organs such as antennae, compound eyes, simple eyes, statocysts (balancing organs), and tactile hairs.
Excretory System:
- Excretion in arthropods occurs through specialized structures called Malpighian tubules.
Reproductive System:
- Arthropods are mostly dioecious (separate sexes), with males and females being distinct individuals.
- Fertilization is typically internal.
- They are mostly oviparous (reproducing by laying eggs).
Development:
- Development in arthropods may be direct (young ones resemble the adult) or indirect (involving larval stages).
Examples:
- Examples of economically important insects include Apis (Honey Bee), Bombyx (Silkworm), and Laccifer (Lac Insect).
- Examples of vectors include Anopheles, Culex, and Aedes (Mosquitoes).
- Examples of gregarious pests include Locusta (Locust).
- Limulus (King Crab) is considered a living fossil.
4.2.8 Phylum – Mollusca
Huge Diversity:
- Mollusca is the second largest animal phylum, comprising a wide variety of species found in terrestrial or aquatic (marine or freshwater) habitats.
Organ-System Level of Organization:
- Molluscs have organ-system level of organization, with distinct organ systems performing specific functions.
Bilateral Symmetry:
- They display bilateral symmetry, with a distinct left and right side.
Triploblastic and Coelomate:
- Molluscs are triploblastic organisms, consisting of three embryonic germ layers: ectoderm, mesoderm, and endoderm.
- They are coelomate animals, possessing a true body cavity (coelom) that is completely lined with mesoderm.
Calcareous Shell:
- The body of molluscs is often covered by a calcareous shell, providing protection and support.
- However, not all molluscs have shells.
Body Structure:
- The body of molluscs is unsegmented and typically consists of three main parts: a distinct head, a muscular foot for locomotion, and a visceral hump containing internal organs.
- A soft and spongy layer of skin called the mantle covers the visceral hump.
Mantle Cavity and Gills:
- The space between the visceral hump and the mantle is called the mantle cavity, which contains feather-like gills.
- The gills serve both respiratory and excretory functions.
Sensory Tentacles:
- Molluscs often have sensory tentacles located on the anterior head region, which aid in detecting food and other environmental stimuli.
Radula:
- The mouth of molluscs contains a file-like rasping organ called the radula, which is used for feeding.
- The radula is unique to molluscs and is adapted for scraping food particles.
Reproductive System:
- Molluscs are usually dioecious (separate sexes) and oviparous (reproducing by laying eggs).
- They often exhibit indirect development, involving larval stages before reaching adulthood.
Examples:
- Examples of molluscs include Pila (Apple Snail), Pinctada (Pearl Oyster), Sepia (Cuttlefish), Loligo (Squid), Octopus (Devil Fish), Aplysia (Seahare), Dentalium (Tusk Shell), and Chaetopleura (Chiton).
4.2.9 Phylum – Echinodermata
Endoskeleton of Calcareous Ossicles:
- Echinoderms have an endoskeleton composed of calcareous ossicles, giving them their characteristic spiny appearance.
Marine Habitat:
- All echinoderms are exclusively marine organisms.
Organ-System Level of Organization:
- Echinoderms exhibit organ-system level of organization, with distinct organ systems performing specific functions.
Radial Symmetry (Adults) and Bilateral Symmetry (Larvae):
- Adult echinoderms typically display radial symmetry, with body parts arranged symmetrically around a central axis.
- However, echinoderm larvae are bilaterally symmetrical.
Triploblastic and Coelomate:
- Echinoderms are triploblastic organisms, consisting of three embryonic germ layers: ectoderm, mesoderm, and endoderm.
- They are coelomate animals, possessing a true body cavity (coelom) completely lined with mesoderm.
Complete Digestive System:
- The digestive system of echinoderms is complete, with a mouth located on the lower (ventral) side and an anus on the upper (dorsal) side.
Water Vascular System:
- The most distinctive feature of echinoderms is the presence of a water vascular system.
- This system consists of a network of fluid-filled canals and tube feet, which are used for locomotion, capturing and transporting food, and respiration.
Absence of Excretory System:
- Echinoderms lack a specialized excretory system.
Separate Sexes and Sexual Reproduction:
- Echinoderms have separate sexes (dioecious), with males and females being distinct individuals.
- Reproduction is typically sexual, with external fertilization.
Indirect Development with Free-Swimming Larvae:
- Development in echinoderms is indirect, involving larval stages before reaching adulthood.
- These larvae are free-swimming and undergo metamorphosis into the adult form.
Examples:
- Examples of echinoderms include Asterias (Starfish), Echinus (Sea Urchin), Antedon (Sea Lily), Cucumaria (Sea Cucumber), and Ophiura (Brittle Star).
4.2.11 Phylum – Chordata
Notochord:
- Presence of a notochord, a flexible rod-like structure derived from mesoderm, providing skeletal support and acting as a precursor to the vertebral column in vertebrates.
- In protochordates, such as Urochordata (Tunicates) and Cephalochordata (Lancelets), the notochord persists throughout life or is present only in the larval stage.
Dorsal Hollow Nerve Cord:
- Presence of a dorsal hollow nerve cord, derived from ectoderm, running along the dorsal side of the body.
- This nerve cord serves as the central nervous system and is a defining feature of chordates.
Paired Pharyngeal Gill Slits:
- Presence of paired pharyngeal gill slits or pouches, which are openings in the pharynx that may function in filter-feeding, respiration, or excretion.
Bilateral Symmetry:
- Chordates exhibit bilateral symmetry, with a distinct left and right side.
Triploblastic and Coelomate:
- Chordates are triploblastic organisms, consisting of three embryonic germ layers: ectoderm, mesoderm, and endoderm.
- They are coelomate animals, possessing a true body cavity (coelom) completely lined with mesoderm.
Post-Anal Tail:
- Presence of a post-anal tail, an extension of the body past the anus, which may be present during embryonic development or persist into adulthood in some species.
Closed Circulatory System:
- Chordates typically have a closed circulatory system, with blood enclosed within vessels and pumped by a heart.
- Subphylum Urochordata (Tunicates) and Cephalochordata (Lancelets) are often referred to as protochordates and are exclusively marine. They exhibit characteristics typical of chordates, such as a notochord, dorsal hollow nerve cord, and pharyngeal gill slits.
- Subphylum Vertebrata encompasses animals with a vertebral column, replacing the notochord during embryonic development. Vertebrates possess additional features such as a ventral muscular heart with chambers, kidneys for excretion and osmoregulation, and paired appendages, which may be fins or limbs.
Examples of chordates include:
- Urochordata: Ascidia, Salpa, Doliolum
- Cephalochordata: Branchiostoma (Amphioxus or Lancelet).
The subphylum Vertebrata is further divided as follows:
1.
Class Agnatha (Jawless Fishes):
- Agnathans are characterized by the absence of jaws.
- They possess a cartilaginous skeleton and typically have a round, eel-like body shape.
- Examples include lampreys and hagfishes.
Class Chondrichthyes (Cartilaginous Fishes):
- Chondrichthyans are characterized by a skeleton made of cartilage instead of bone.
- They typically have paired fins, a streamlined body, and placoid scales.
- Examples include sharks, rays, and skates.
Class Osteichthyes (Bony Fishes):
- Osteichthyans are characterized by a skeleton made of bone.
- They have gills covered by a bony operculum and typically possess swim bladders for buoyancy control.
- Examples include ray-finned fishes (Actinopterygii) and lobe-finned fishes (Sarcopterygii).
Class Amphibia (Amphibians):
- Amphibians are tetrapod vertebrates that typically undergo metamorphosis from an aquatic larval stage to a terrestrial adult stage.
- They have moist skin, lay eggs in water or damp environments, and often have a biphasic life cycle.
- Examples include frogs, toads, salamanders, and caecilians.
Class Reptilia (Reptiles):
- Reptiles are tetrapod vertebrates that are characterized by scales or scutes covering their body.
- They are ectothermic (cold-blooded) and lay shelled eggs on land.
- Examples include snakes, lizards, turtles, crocodilians, and tuatara.
Class Aves (Birds):
- Birds are characterized by feathers, a beak without teeth, and a lightweight skeleton with pneumatic bones.
- They are endothermic (warm-blooded) and possess adaptations for flight.
- Examples include eagles, sparrows, penguins, and ostriches.
Class Mammalia (Mammals):
- Mammals are characterized by mammary glands, hair or fur, and a neocortex region in the brain.
- They are endothermic (warm-blooded) and typically give birth to live young.
- Examples include humans, dogs, cats, whales, and elephant.